Rice LC2 (OsVIL3) and OsVIL2 (two OsVILs, possible components of PRC2 complex) promote rice flowering by binding to the promoter region of a floral repressor OsLF and represses the OsLF expression via H3K27 tri-methylation modification. OsLF directly regulates the Hd1 expression through binding to Hd1 promoter.